300 miles isn't too far for spiders to float across. the idea that highly specialized, hard-to-find, or "very unique" animals are less likely to develop on the mainland doesn't make any sense. because nobody is calling this convergent evolution, this type of spider presumably -did- evolve more than 165 mya, which isn't that shocking.Madagascar is home to some of the world's most unique flora and fauna. Almost all of Madagascar's reptile and amphibian species, half of its birds, and all of its lemurs are endemic to the island. (they exist nowhere else on earth). so that is why it doesnt surprise me. even tho south africa is only 300 miles west of madagascar it surprises me because it (madagscar) has been cut off, unexplored for so many years, and the fact that something as unique and highly specialised as this spider has evolved in such a place as south africa and australia. basicly, it surprises me for the exact opposite reasons it doesnt surprise me it was found in madagascar. (if that makes sense)
this means that the assasin spiders have either evolved the same way as on madagascar seperately or they have been around for more than 165 million years. because madagascar was attached to africa more than 165 million years ago. However, most of the groups of mammals and other terrestrial fauna that are well represented on Madagascar today had not evolved when Madagascar first split from continental Africa.
i hope this answered your question