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- Dec 18, 2018
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So a few days ago on this forum I had a discussion about Emperor Scorpion sizes. This caused me to go into a bit of a deep dive, and since then I've been going into an extremely deep dive into Emperor scorpion sizes and their reported size ranges. This is what I have so far. This information is partially incomplete, a lot of the earlier sources I couldn't find, and/or are not written in English. If anyone has any more information to add to this thread, I would wholly appreciate it.
The Timeline:
1841: Koch describes Pandinus Imperator. The holotype was rediscovered and redescribed in 2017. Though the specimen measured 160mm, it might've been 170mm accounting for some damage.
1956: The Biology of Scorpions by Max Vachon. Only reference to Pandinus imperator size was in a drawing, with a stated range of 18-20cm. This range is cited as "After Gaillard", but I could not find what this was referring to. No specific measurements of any individual given.
1980: Scorpions of Medical Importance by Hugh L. Keegan. Size for Pandinus imperator stated as "up to 180mm or more".
1985: Prey capture and stinging behavior in the emperor scorpion, Pandinus imperator by Gary S. Casper. First source with reported measurements of specimens. Largest measured at 17cm from chelicerae to telson tip, numbers likely rounded as the scorpions were sorted into groups of size, no specific measurements of any individual.
1988: How to Keep Scorpions by Hull-Williams. Conflicting information, says scorpions (no specific species or family) get "between 40mm and 127mm", then later in the text says "Large forest dwelling species are nocturnal ramblers and also grow quite large (8-9")". In the section on Heterometrus longimannus says "Due to their conspicuous size (trunk length up to 160mm) they are a good match for the closely related African genus Pandinus Thorell: they are among the world's largest scorpions." Does not give a size range for Pandinus imperator.
1990: The Biology of Scorpions by Gary A. Polis. Given range for Pandinus imperator is 18-20cm and references "Vachon, 1953" for this range. No specific reference as to what actual paper or publication this is.
1991: Arachnomania by Philippe De Vosjoli. Says Pandinus imperator "reach a length of nearly eight inches (20cm)", references include Scorpions of Medical Importance by Hugh L. Keegan, The Biology of Scorpions by Gary A. Polis, and How to Keep Scorpions by Hull-Williams. We can infer the 20cm figure is from Gary A. Polis, who in turn cited Vachon for this.
1994?: CITIES proposal to include multiple Pandinus species, including Pandinus imperator. Says "reaching as much as 20cm", but doesn't cite where this specific number is from. Includes a paper by Vachon in 1967 in the reference section, though I could not find this paper and it's not in English.
1999: Variation in energy spent on reproduction between forest and savanna populations of Pandinus imperator (Koch) in the Ivory Coast by LOURENCO,W.R. & CLOUDSLEY-THOMPSON,J.L. First source with specific measurements of individuals, categorizes the difference in sizes between larger "forest" form Emperors and smaller "savannah" form Emperors. Unfortunately does not give a table with their precise measurements, and only measure the prosoma + mesosoma, without including the metasoma. Largest individual measures appears to be about 82mm, assuming the metasoma is not significantly longer than the prosoma + mesosoma gives us a length of about 164mm.
2010: Scorpions of the World by Roland Stockmann and Eric Ythier. Given size range is up to 200mm.
2011: A Review of the Subgenus Pandinus Thorell, 1876 with Descriptions of Two New Species from Uganda and Ethiopia by František Коvařík. Stated size range of Pandinus imperator given as 220mm (the largest so far), does not include a material examined section. Only one measurement of a specific specimen given, a male from Ghana reported to be 115mm.
2014: Notes on the distribution of Pandinus (Pandinus) Thorell, 1876 and Pandinus (Pandinurus) Fet, 1997 with the descriptions of two new species from Central African Republic and Djibouti by Andrea Rossi. Size range reported as up to 230mm (the largest estimate given to date). and does include a materials examined section. Specimens examined are one female from Togo/Ghana (ARPC) later measured to be 113mm, two males and two females from Liberia (MHNG), with one male stated as being "about 180mm". and one male "without data" (MSNB).
2014: Further considerations on the identity and distribution of Pandinus imperator (C. L. Koch, 1841) and description of a new species from Cameroon by WILSON R. LOURENÇO. Reported size range for Pandinus imperator up to 180mm, gives specific values for a male from SE Notse, South of Togo (154.8mm) and a female Edea South of Cameroon (163.3mm). The female Pandinus imperator was suspected to be a P dictator by a later paper.
2015: Clarification of the Type Locality of Pandinus Ulderigoi with Notes on the Scorpions Protected by CITES by Andrea Rossi. No specific size range for Pandinus imperator given, again shows the same picture of the male from Liberia (MHNG) captioned as being 180mm. The male "without data" (MSNB) is no longer listed on this paper, though the 4 specimens from Liberia (MHNG) and a male from the Ivory Coast (ARPC) are listed.
2016: Redefinition and Systematic Revision of the East African Scorpion Genus Pandinoides (Scorpiones: Scorpionidae) With Critique of the Taxonomy of Pandinus, Sensu Lato by Lorenzo Prendini. Doesn't give specific size ranges but important to note as he also talks about the differences in sizes of Pandinus imperator based on locality.
2023: A new species of Pandinus Thorell, 1876 from the Sahelian wooded steppes of Burkina Faso by Ythier, Eric & Audibert, Cédric. Gives the size range as up to 230mm, though this was likely taken from Rossi's 2014 paper as the authors explicitly note they were examining specimens from the smaller localities.
2017 Rediscovery of Holotype
1956 Vachon
1985 Casper
1994 CITIES
1999 Lourenco & Cloudsley-Thompson
2011 Kovarik
2014 Rossi
2014 Lourenco
2015 Rossi
2016 Prendini
2023 Ythier & Audibert
I do not have digital copies of The Biology of Scorpions by Gary A. Polis, Arachnomania by Philippe De Vosjoli, Scorpions of Medical Importance by Hugh L. Keegan, Scorpions of the World by Stockmann & Ythier, or How to Keep Scorpions by Hull-Williams.
To summarize, it seems 20cm has been the reported max size of Emperors for a while, but no specific specimen with these measurements has been reported from what I could find. The number likely traces back to Vachon, who in the one source I was able to find cited the number to Gaillard. I would like to see more of these sources if possible to see if such a specimen was ever actually recorded, and see how precisely they were measuring it. The 8-9in number from How to Keep Scorpions likely included the pedipalps, as that number is much larger than the 160mm later stated in the book. Kovarik is the first source to say anything above 200mm, but he also doesn't reference a specific specimen he measured at 220mm. Rossi has the largest number thus far, but again does not give the measurements of the specific individual he measured at 230mm. I assume it was the male "without data" (MSNB), and this specimen is also no longer cited in the paper he published the next year. The largest scorpion we have a picture of for having physically existed is the male from Liberia in Rossi's 2014 and 2015 paper, initially stated to be "about 180mm" then later just stated to be "180mm", and though the 2023 paper by Ythier and Audibert also give the range as 230mm, I believe it is likely they were going off of Rossi's 2014 paper.
So it seems from this, the largest scorpions we actually measure seem to top out at 170mm, with the potential for them to get up to 180mm if Rossi's numbers are accurate for the specimen from Liberia. This seems to be an extremely uncommon occurrence though, and even then these sizes would only be obtainable in the larger "forest" form Emperors, which are incredibly uncommon in the hobby, especially in recent years. The smaller "savannah" form Emperors seem to get between 100-130mm, which is consistent with the Emperors I've seen both personally and posted online.
If anyone has more information, especially any evidence of a scorpion getting to or exceeding 180mm, I would greatly appreciate it. If anyone has any links to some of Vachon's other papers, or has any idea what "Gaillard" refers to, I would also appreciate the insights. I am also interested as to what the largest scorpion we actually have on record is, even in G swammerdami 200mm+ does not seem very common, if possible. Let me know your thoughts.
The Timeline:
1841: Koch describes Pandinus Imperator. The holotype was rediscovered and redescribed in 2017. Though the specimen measured 160mm, it might've been 170mm accounting for some damage.
1956: The Biology of Scorpions by Max Vachon. Only reference to Pandinus imperator size was in a drawing, with a stated range of 18-20cm. This range is cited as "After Gaillard", but I could not find what this was referring to. No specific measurements of any individual given.
1980: Scorpions of Medical Importance by Hugh L. Keegan. Size for Pandinus imperator stated as "up to 180mm or more".
1985: Prey capture and stinging behavior in the emperor scorpion, Pandinus imperator by Gary S. Casper. First source with reported measurements of specimens. Largest measured at 17cm from chelicerae to telson tip, numbers likely rounded as the scorpions were sorted into groups of size, no specific measurements of any individual.
1988: How to Keep Scorpions by Hull-Williams. Conflicting information, says scorpions (no specific species or family) get "between 40mm and 127mm", then later in the text says "Large forest dwelling species are nocturnal ramblers and also grow quite large (8-9")". In the section on Heterometrus longimannus says "Due to their conspicuous size (trunk length up to 160mm) they are a good match for the closely related African genus Pandinus Thorell: they are among the world's largest scorpions." Does not give a size range for Pandinus imperator.
1990: The Biology of Scorpions by Gary A. Polis. Given range for Pandinus imperator is 18-20cm and references "Vachon, 1953" for this range. No specific reference as to what actual paper or publication this is.
1991: Arachnomania by Philippe De Vosjoli. Says Pandinus imperator "reach a length of nearly eight inches (20cm)", references include Scorpions of Medical Importance by Hugh L. Keegan, The Biology of Scorpions by Gary A. Polis, and How to Keep Scorpions by Hull-Williams. We can infer the 20cm figure is from Gary A. Polis, who in turn cited Vachon for this.
1994?: CITIES proposal to include multiple Pandinus species, including Pandinus imperator. Says "reaching as much as 20cm", but doesn't cite where this specific number is from. Includes a paper by Vachon in 1967 in the reference section, though I could not find this paper and it's not in English.
1999: Variation in energy spent on reproduction between forest and savanna populations of Pandinus imperator (Koch) in the Ivory Coast by LOURENCO,W.R. & CLOUDSLEY-THOMPSON,J.L. First source with specific measurements of individuals, categorizes the difference in sizes between larger "forest" form Emperors and smaller "savannah" form Emperors. Unfortunately does not give a table with their precise measurements, and only measure the prosoma + mesosoma, without including the metasoma. Largest individual measures appears to be about 82mm, assuming the metasoma is not significantly longer than the prosoma + mesosoma gives us a length of about 164mm.
2010: Scorpions of the World by Roland Stockmann and Eric Ythier. Given size range is up to 200mm.
2011: A Review of the Subgenus Pandinus Thorell, 1876 with Descriptions of Two New Species from Uganda and Ethiopia by František Коvařík. Stated size range of Pandinus imperator given as 220mm (the largest so far), does not include a material examined section. Only one measurement of a specific specimen given, a male from Ghana reported to be 115mm.
2014: Notes on the distribution of Pandinus (Pandinus) Thorell, 1876 and Pandinus (Pandinurus) Fet, 1997 with the descriptions of two new species from Central African Republic and Djibouti by Andrea Rossi. Size range reported as up to 230mm (the largest estimate given to date). and does include a materials examined section. Specimens examined are one female from Togo/Ghana (ARPC) later measured to be 113mm, two males and two females from Liberia (MHNG), with one male stated as being "about 180mm". and one male "without data" (MSNB).
2014: Further considerations on the identity and distribution of Pandinus imperator (C. L. Koch, 1841) and description of a new species from Cameroon by WILSON R. LOURENÇO. Reported size range for Pandinus imperator up to 180mm, gives specific values for a male from SE Notse, South of Togo (154.8mm) and a female Edea South of Cameroon (163.3mm). The female Pandinus imperator was suspected to be a P dictator by a later paper.
2015: Clarification of the Type Locality of Pandinus Ulderigoi with Notes on the Scorpions Protected by CITES by Andrea Rossi. No specific size range for Pandinus imperator given, again shows the same picture of the male from Liberia (MHNG) captioned as being 180mm. The male "without data" (MSNB) is no longer listed on this paper, though the 4 specimens from Liberia (MHNG) and a male from the Ivory Coast (ARPC) are listed.
2016: Redefinition and Systematic Revision of the East African Scorpion Genus Pandinoides (Scorpiones: Scorpionidae) With Critique of the Taxonomy of Pandinus, Sensu Lato by Lorenzo Prendini. Doesn't give specific size ranges but important to note as he also talks about the differences in sizes of Pandinus imperator based on locality.
2023: A new species of Pandinus Thorell, 1876 from the Sahelian wooded steppes of Burkina Faso by Ythier, Eric & Audibert, Cédric. Gives the size range as up to 230mm, though this was likely taken from Rossi's 2014 paper as the authors explicitly note they were examining specimens from the smaller localities.
2017 Rediscovery of Holotype
1956 Vachon
1985 Casper
1994 CITIES
1999 Lourenco & Cloudsley-Thompson
2011 Kovarik
2014 Rossi
2014 Lourenco
2015 Rossi
2016 Prendini
2023 Ythier & Audibert
I do not have digital copies of The Biology of Scorpions by Gary A. Polis, Arachnomania by Philippe De Vosjoli, Scorpions of Medical Importance by Hugh L. Keegan, Scorpions of the World by Stockmann & Ythier, or How to Keep Scorpions by Hull-Williams.
To summarize, it seems 20cm has been the reported max size of Emperors for a while, but no specific specimen with these measurements has been reported from what I could find. The number likely traces back to Vachon, who in the one source I was able to find cited the number to Gaillard. I would like to see more of these sources if possible to see if such a specimen was ever actually recorded, and see how precisely they were measuring it. The 8-9in number from How to Keep Scorpions likely included the pedipalps, as that number is much larger than the 160mm later stated in the book. Kovarik is the first source to say anything above 200mm, but he also doesn't reference a specific specimen he measured at 220mm. Rossi has the largest number thus far, but again does not give the measurements of the specific individual he measured at 230mm. I assume it was the male "without data" (MSNB), and this specimen is also no longer cited in the paper he published the next year. The largest scorpion we have a picture of for having physically existed is the male from Liberia in Rossi's 2014 and 2015 paper, initially stated to be "about 180mm" then later just stated to be "180mm", and though the 2023 paper by Ythier and Audibert also give the range as 230mm, I believe it is likely they were going off of Rossi's 2014 paper.
So it seems from this, the largest scorpions we actually measure seem to top out at 170mm, with the potential for them to get up to 180mm if Rossi's numbers are accurate for the specimen from Liberia. This seems to be an extremely uncommon occurrence though, and even then these sizes would only be obtainable in the larger "forest" form Emperors, which are incredibly uncommon in the hobby, especially in recent years. The smaller "savannah" form Emperors seem to get between 100-130mm, which is consistent with the Emperors I've seen both personally and posted online.
If anyone has more information, especially any evidence of a scorpion getting to or exceeding 180mm, I would greatly appreciate it. If anyone has any links to some of Vachon's other papers, or has any idea what "Gaillard" refers to, I would also appreciate the insights. I am also interested as to what the largest scorpion we actually have on record is, even in G swammerdami 200mm+ does not seem very common, if possible. Let me know your thoughts.
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